i raced a few seasons with VM26s, they are actuallly 27mm ID, relatively cheap (compared to VM28s), easy to setup and bike was definitely fast. went with CR29s later, difference "might" have been at the very top end, maybe, never got around to dyno it with the CRs for a comparison. the 500/550 are so restricted in flow in the valve throat area that i find it hard to believe that you will feel much difference in street riding.
my 0.02$