The 550 has 1.85 inch rims front and rear.
The tire manufacturers often state the allowable rim size that works with the selected tire if you look hard enough. Tire salesmen hide this info from you as there is more profit margin is larger tires. Plus, fitting them onto narrow rims helps them wear faster, and has you coming back for replacements sooner. Why? Because the rim is pinched together making a higher sidewall angle that has less rigidity and adding more "crown" the tread area at the same time. To stiffen the sidewall, more tire pressure is required and that adds even more crown to the tire tread, resulting in much more rapid wear in the center of the tire tread.
I don't let them sell me metric "equivalents" anymore. Because they aren't really equivalent.
The old inch size tires for the 550 were/are 100% aspect ratio. A 3.25 front equates to 83 mm width and height. A 90/90 tire is wider and shorter at only 81mm height. The good news is that it will usually work ok for a 1.85 inch width rim. But, it works best with a wider rim of 2.15 inch. And, bikes designed to use metric tires have wider rims than the 550.
A 3.75 rear equates to 95mm width and height. A 100/90 width tire is wider and shorter, and is optimized for an even wider rim than the 90/90.
Actually a 4.00/18 rear is far easier to get and has a 102mm equivalent width. But, to get close to that height, you have to go to a 110/90 (99mm height) which works best on a 2.50 rim width that the 550 doesn't have.
A 120 metric tire is simply too wide for the 550s 1.85 inch rim width. It will center wear far faster than the proper tire, and require much earlier replacement.
That's why I still use inch size tires, which the tire hawker will order for you if you insist. But, they will try their best to sell from what they stock at the store using what are convincing arguments for the uneducated.
Finally, I think the the 550 simply handles better with the proper inch size tires.
Cheers,
See also:
http://www.dkebay.com/jsp/tpl/tire/tiresizeexplain.jsp