Just like you can fit a tubeless style tire onto a spoked wheel, you can also put them on Comstars, as well. Still should use an inner tube, unless the rim states it is designed to use the tubeless type, or the rim manufacturer says it is correct fitment.
I believe the issue is seating area of tire to rim. The seating area width is sized, and wheel diameter tolerance is maintained, to provide a positive seal between rim and tire bead, in order to give reliable air retention, and bead stability, including when the tire pressures are very low. (slow leak, puncture, etc.)
Non-tubeless rims don't have the bead seal requirement, so those size and tolerance requirements are hit or miss and the air seal function is the sole responsibility of the inner tube.
I really can't speak for bikes outside of the US, which may state different requirements in their owner's manuals. But, it does seem odd that Honda would provide wheel differences depending on export type, unless there were government requirement issues or other unknown cost/supply factors.
I'm not arguing that the tubeless system isn't the better way to go. It's just that the benefits of tubeless aren't there by providing half the requirements for the tubeless system. The two halves are tire AND rim designs.
Cheers,