At this point, run it and see. I replace them on 400F and 350F engines because they are usually plasticized if they are original. I have no idea what they seal, and can't see what they clamp to, but I assume they are there for a reason. Maybe to seal against leaks between the sleeve and the jug?
For what it is worth, the originals definitely sit flush with the lower clamping surface, so it looks like they contact something. I say this because the new rings sit proud of the clamping surface, which leads me to believe that they contact something to create a seal.
Camelman